In a gritty fantasy world where dinosaurs did not become extinct, human history has unfolded side-by-side with the giant reptiles. The greatest of the dinosaur slayers are known as the Predator Defense League. When an act of kindness causes thousands of the beasts to destroy the walls of Monanti City, it leaves culture vs. culture, mammal vs. reptile, friend vs. friend and father vs. daughter.
Lilli and Pax escape the dinosaur hoard into an ancient village at the tops of the trees where they find a clue to unlocking the mystery of the little girl and the reptile attacks. Meanwhile, the scattered Predator Defense League looks to find allies in their fight against the Talpid occupation.