The rise of the Tzalekuhl Empire threatens to disrupt the peace that has lasted for generations across the land of Mezo. When the conquest begins, a young girl named Kyma witnesses the death of her father, Hegol, a tribal leader who refused to yield. As the solar eclipse nears, Kyma must unite the various tribes against an emperor determined to make them all kneel before his god or be sacrificed.
Emperor Vuh presents his sacrfice to the gods while Roden watches on with his family. With the help of Ne'nezi, they are able to extract the location of the Huax'kin so they can hunt them down and finish the job. Meanwhile, Kyma has led her tribe to the Kan'kin in hopes of convincing them to unite against a common enemy.