The rise of the Tzalekuhl Empire threatens to disrupt the peace that has lasted for generations across the land of Mezo. When the conquest begins, a young girl named Kyma witnesses the death of her father, Hegol, a tribal leader who refused to yield. As the solar eclipse nears, Kyma must unite the various tribes against an emperor determined to make them all kneel before his god or be sacrificed.
The great Shaman, Itzabé, returns to land of Mezo in hopes of stopping the Talekuhl conquest. He's brought with him a young protege named Zea and the two of them make friends with twin warriors on their quest for the World Tree. Meanwhile, Roden's decisive actions in battle have earned him a promotion to Palatzin in the Tzalekuhl army.